I'm pretty sure you are seeing an event called "Red Plating" which is the tube overheating. There are a few causes- and finding them can be a mystery.
The tube could be shorting (doubtful given what you've described)
The bias is certainly off. How could I know that? All biases are off at least a little. Add time, other component deterioration, and mismatched components and it's a boas nightmare.
I had a mystery red plating problem and it turned out to be a bad 12BH7 high current driver - which caused me to have to replace 2 6550s. OUCH!