@willmarchant , I may be missing something, but that is interesting that you would have a tube with no bias reading but it would still sound good?
When my Cary amplifier went into a bizarre biasing mode on one channel, one of the things I was advised (not by Cary) to do was to take my MM and take readings on everything in the chassis and see if anything was wildly different than it's partner. Can you get your probes on those resistors that ARC was referring to?
(On mine, my MM identified a signal cap that read way different than the other three. Under further advice (again, not from Cary) I bought a cap checker and confirmed that the cap was bad. I replaced the cap, and to my surprise, but delight, that solved the bizarre biasing problem. So I went ahead & replaced the other 3 caps.)